Karlo graduated at the Faculty of Science, University of Zagreb, in 2019 with a Master's degree in Molecular Biology with magna cum laude honors. He was working on his master's thesis project in Plant Tissue Culture Laboratory under the mentorship of Assoc. prof. Dunja Leljak-Levanić where he focused on BPM1 protein as a novel regulator of RNA directed DNA methylation in plant develompent and its importance in de novo methylation of plant genome, using bacteria, yeast and plant models. Karlo just joined Zoldos Lab and is interested in genomic engineering and regulation of genes involved in imunoglobulin G glycosylation. He is using CRISPR/dCas9 technology and HEK293FreeStyle transient system for IgG production with a focus on glycoengineering of IgG antibody within the CRISPR/dCas9 - CasMouse project.